Telephone: 44 (0) 1631 559233
   E-mail: maeve.kelly@sams.ac.uk

Scientific interests:

My research focuses on the reproductive and larval biology, growth, nutrition and ecology of commercially important marine invertebrates. I also research the occurrence, distribution and biological implications of harmful algal-toxin accumulation in shellfish. Linking these themes is my interest in integrated aquaculture, polyculture, sustainability and bioremediation in marine systems. I have strong links with academic and industrial partners in a variety of disciplines and at international, European, national and local level. I have secured the funding for and co-ordinated projects at European, National and local level. I am module leader for the SAMS UHI Marine Science degree’s Level 3 Aquaculture Module. I have over 40 fully peer-reviewed publications in international journals.


Current Projects / Grants:
• 2007 SAMS component of SUPERGEN consortium bid ‘Potential of marine biomass to UK energy, fuels and chemicals. Value to SAMS £80,000. Start date June 2007. 4 years duration. www.supergen-bioenergy.net

• 2007 MARFUEL: The potential of Marine Biomass as Biofuel. A feasibility study with research recommendations for further research. Funded by The Crown Estate. £30,000. Start date Jan 2007.

• 2007 AAAG2 Atlantic Arc Aquaculture Group (funded by ERDF Interreg IIIB)– project renewal – value to SAMS 187,050 Euros. Start date Jan 2007. 18 months duration, Integrated Aquaculture Research www.arcaqua.org

• 2006 MARFEED: Alternative marine sources for replacement fish oil and fish meal. A literature review with recommendations for future research avenues. Funded by The Crown Estate. £23,000. Start date Nov 2005.

• SPIES-DETOX. An EU funded collective grant Active biological monitoring and removal of toxins in aquaculture ecosystems and shellfish - including the development of a Solid-Phase In-situ Ecosystem Sampler (SPIES) and detoxification of shellfish (DETOX). Proposal/Contract no.: 030270-2. Start date July 1st 2006. Value to SAMS 153,696 Euros.

• 2006 COMMWEED – Commercialising seaweed culture in conjunction with Atlantic salmon. A Highlands and Islands funded initiative to further seaweed cultivation. Value to SAMS £45,000. Start date 1st August 2006. 18 months duration.

• 2005 Larval Invertebrate Microalgal Interactions. A FP6 Mobility Marie Curie award for Incoming International Fellows (Prof. Hui Liu, Chinese Academy of Fishery Science). Value €135,425. Start date June 2005, I year duration.

• 2004 MERMAIDS: Macroalgae for Environmental Remediation, Managing Aquaculture Initiatives for Diversification and Sustainability. A SAMS/ Western Isles based AIE funded PhD studentship (Mr Andrew Rodger). £36k. Start date October 2004

• 2004 SPIINES 2: Sea urchin Production In Integrated systems, their Nutrition and roe Enhancement. An EC funded CRAFT project. Start date January 2005, for 27 months. RTD Co-ordinator MSK, total project value €970,640; value to SAMS €183,860. www.spiines.com

• 2004 KEYZONES: To investigate sustainable biological carrying capacities of Key European coastal zones. An EC funded CRAFT project. Start date January 2005. Workpackage 2 led by SAMS, value to SAMS €158922. www.keyzones.com

• 2003 Atlantic Arc Aquaculture Group (funded by ERDF Interreg IIIB) December 2003 - 2006. A collaborative project led by University of North wales with project partners in France, Spain and Ireland. SAMS research themes include sustainablility and bioremediation in aquaculture. Value to SAMS €860,000. www.arcaqua.org

• 2003 REDWEED: Reducing the environmental impact of sea cage fish farming through the cultivation of seaweeds. A SAMS-UHI PhD Studentship (Mr Craig Sanderson). £114k. Start date February 2003. (funded by Highland Council and HIE).

• 2003 ALGINET: an EU thematic network; Microalgae as Cell Factories for Chemical and Biochemical Products value to SAMS €21,600 over three years. Start date March 2003.

• 2002 TALISMAN: Algal toxins, their accumulation and loss in commercially important shellfish, including larval mortality and appraisal of accelerated depuration. An EC funded CRAFT project, December 2002 – April 2005. Value to SAMS £120k.

• 2002 ‘Sea bed biota, scraper and shapers, urchins in control?’ A SAMS-UHI NERC funded PhD studentship (Mr Adam Hughes). Start date November 2002.

Previous Research Grants:
• 2001 Highland Council and HIE. Tube worm fouling on rope grown mussels – its importance to the industry. £41k

• 2000 – 2003. A SAMS-UHI funded PhD Diets, energetics and ecology of the sea urchin Psammechinus miliaris. UHI Millennium Institute (Maria del Mar Otero Villanueva).

• 2000 Highland Council. Designing and testing cost effective finishing diets for the sea urchin Psammechinus miliaris. £5.2k

• 1998 - 2000 EC CRAFT Co-operative Award: Commercially Viable Sea Urchin Culture £390k

• 1996 - 1999 LINK Aquaculture Programme: Echinoculture: Studies on farming the green sea urchin Psammechinus miliaris £340.78k (including 50% in kind from industry)

• Dec 1999 – May 2000 HIE, Highland Council, PESCA and the Scallop Association: Occurrence and distribution of the ASP toxin in king scallop population on the west coast of Scotland. £34.2k

• Jun 2000 – Dec 2000 HIE, Highland Council and the Scallop Association. ASP in the King Scallop, Pectin maximus, from the West Coast of Scotland. £31k

• 1999 – 2000 Argyll and Islands Enterprise and Single Shell Systems: Accelerating growth rates in the King Scallop Pecten maximus £40.2k.

• 1998 – 1999 Lochaber PESCA Project and The Highland Council: Abalone Farming in Scotland – A preliminary study £24k

• 1998 – 1999 Highlands and Islands Enterprise and Highland Council: Echinus esculentus: the feasibility of designing an artificial diet to enhance roe quality. £30k

• 1997 EC CRAFT Scheme Exploratory Award: Commercially Viable Sea Urchin Culture £37k

• 1996 Highlands and Islands Enterprise: Manufacture of an Artificial Diets for the green sea urchin Psammechinus miliaris from cast Macroalgal fronds £30k

• 1995 Highland and Islands Enterprise: Psammechinus miliaris; its potential as a polyculture species in Scotland (with JDMcK) £18k

• 1992, Waitengi Fellowship, Bank of Scotland and Countrywide Bank, New Zealand. £4.5k

Teaching / supervising activities
SAMS UHI Marine Science degree’s Level 3 Aquaculture Module

Publications Submitted:
1. Carotenoids in the sea urchin Paracentrotus lividus: occurrence of 9’-cis echinenone as the dominat carotenoid in gonad colour determination. Symonds, R.C., Caris-Veyrat, C., Kelly, M.S., Young, A.J. Comp. Biochem. Physiol.

2. Liu H., Kelly, M.S., Campbell, D.A., Dong, S.L., Zhu, J. X., Fang, J.G. and Wang, S. F. Exposure to domoic acid affects juveniles of the king scallop Pecten maximus. Harmful Algae.

Publications 2007:
1. Comparative study of the antiinflammatory, anticoagulant, antiangiogenic, and antiadhesive activities of nine different fucoidans from brown seaweeds. Cumashi1 A.,. Ushakova2 N. A.,, Preobrazhenskaya2 M. E., D’Incecco1 A., Piccoli3 A.,, Totani,3 L., Tinari1 N., Morozevich2 G.E., Berman2 A. E., Bilan4 M. I., Usov4 A. I., Ustuzhanina5 N. E., Sanderson6 C. J., Kelly6 M. , Rabinovich7 G. A., Iacobelli1 S., and Nifantiev5,8 N., E. Glycobiology. Available on line Feb 2007.

2. Cook, E.J. and Kelly M.S. 2007. Effect of variation in the protein value of the red macroalga Palmaria palmata on the feeding, growth and gonad composition of the sea urchins Psammechinus miliaris and Paracentrotus lividus (Echinodermata). Aquaculture. Corrected proof available on line.

3. Liu H., Kelly, M.S., Campbell, D.A., Dong, S.L., Zhu, J. X., Fang, J.G. and Wang, S. F. 2007. Exposure to domoic acid affects larval development of the king scallop Pecten maximus. Aquatic Toxicology 81 152-158.

4. Liu, H., Kelly, M.S., Campbell, D.A., Dong, S.L., Zhu, J.X., Wang, S.F., 2007. Ingestion of Domoic Acid and its Impact on King scallop Pecten maximus (Linnaeus, 1758). Journal of Ocean University of China, in press.

5. Liu H., Kelly M.S., Cook E.J., Black, K.D., Orr H., Zhu, J.X. and Dong, S.L. 2007. The effect of diet type on growth and fatty-acid composition of sea urchin larvae, II Paracentrotus lividus (Lamarck, 1816) (Echinodermata). Aquaculture 264: 247 – 262

6. Liu H., Kelly M.S., Cook E.J., Black, K.D., Orr H., Zhu, J.X. and Dong, S.L. 2007. The effect of diet type on growth and fatty-acid composition of sea urchin larvae, I Psammechinus miliaris (Gmelin). Aquaculture 264: 263 – 278.

7. Robson, A.A., Kelly, M.S. and Latchford, J.W. 2007. Effect of temperature on the spoilage rate of whole, unprocessed crabs: Carcinas maenas, Necora puber and Cancer pagurus. Food Microbiology 24 (4) 419-424.

8. Kelly, M.S., Hughes A.D. and Cook, E. J. 2007. The ecology of Psammechinus miliaris. In The Biology and Ecology of Edible Echinoid Species 2nd Edition (J. Lawrence, Editor) Elsevier Science, Amsterdam.

Peer Reviewed Publications:
9. Wilding TA, Kelly MS, Black KD. 2006. Alternative marine sources of protein and oil for aquaculture feeds: State of the art and recommendation for further research. The Crown Estate, ISBN 978-0-9553427-4-5.

10. Otero-Villanueva, M., Kelly, M.S. and Burnell, G. 2006. Foraging decisions by the regular echinoid Psammechinus miliaris. Journal of the Marine Biological Association, UK. 86, 773-781

11. Hughes, A. D., Kelly, M. S., Barnes, D. K. A., Catarino A., and Black K. D. 2006. The dual function of sea urchin gonads are reflected in the temporal variations of their biochemistry. Marine Biology 148, 4: 789 – 798

12. Kelly, M.S. 2005. Psammechinus miliaris CAB International, [original text by MS Kelly]. In: Aquaculture Compendium. Wallingford, UK

13. Kelly, M.S. 2005. Echinus esculentus CAB International, [original text by MS Kelly]. In: Aquaculture Compendium. Wallingford, UK

14. Hughes, A. D., Catarino A., Kelly, M. S., Barnes, D. K. A and Black K. D. 2005. Gonad Fatty Acids and Trophic Interactions: why living on the edge pays for the echinoid Psammechinus miliaris. Marine Ecology Progress Series 305: 101-111

15. Kelly, M.S. 2005. Book Chapter ‘Echinoderms: their culture and bioactive compounds’. In Progress in Molecular and Subcellular Biology; Subseries Marine Molecular Biotechnology Ed V. Matranga. (series editor W.E.G.Muller) Springer-Verlag, Berlin, Heidelberg, pp 140 – 165. ISBN

16. Kelly, M.S. 2004. Sea Urchin Fisheries and Aquaculture: A Review and Outlook. Echinoderms: Munchen. (Proceedings of the 11th International Echinoderm Conference, Munich, 2003). Eds Heinzeller, T and Nebelsick, J.H. pp 283 – 289. ISBN 04 1536481 7.

17. Kelly, M.S. 2004. Echinus esculentus, a fully farmed approach for a designer roe? In J Lawrence and O. Guzman (eds) Sea urchins: fisheries and aquaculture. Pennsylvania, DEStech publications 256-263.

18. Otero-Villanueva, M., Kelly, M.S. and Burnell, G. 2004. How diet influences energy partitioning in the regular echinoid Psammechinus miliaris; constructing an energy budget. Journal of Experimental Marine Biology and Ecology 304(2): 159-181.

19. Campbell, D. A., Kelly, M.S., Busman, M., Wiggins, E., and Fernandes T.F. 2003. Impact of preparation method on gonad domoic acid levels in the scallop, Pecten maximus (L.) HARMFUL ALGAE (2) 215 – 222.

20. Otero, M., Kelly, M.S. and Burnell, G.M. 2003. Feeding Behavour of Psammechinus miliaris, a dominant grazer of invertebrate communities. Proceedings of the 6th European Conference on Echinoderms, Banyuls, France. Balkema, Rotterdam, pp 249 – 252.

21. Jimmy, R.A, Kelly, M.S. and Beaumont, A.R. 2003. The effect of diet type and quantity on the development of common sea urchin larvae Echinus esculentus. Aquaculture 220 (1-4): 261-275

22. Kelly, M.S. 2002. Survivorship and growth rates of hatchery-reared sea urchins. Aquaculture International. 10 (4): 309-316 2002

23. Kelly, M.S. and Owen, P.V. 2002. Effects of Temperature and Diet on the Growth of the Abalone Haliotis tuberculata L. at Scottish sea temperatures. Aquaculture Research 33: 729 – 733

24. Campbell, D.A. and Kelly, M.S. 2002. Settlement of Pomatoceros triqueter (L.) in two Scottish lochs, and factors determining its abundance on mussels grown in suspended culture. Journal of Shellfish Research 21 (2) 519 – 527

25. Kelly, M.S., Owen, P.V. and Pantazis, P. 2001. The commercial potential of the common sea urchin Echinus esculentus from the west coast of Scotland. "Coastal shellfish – a sustainable resource" Editor Gavin Burnell, Proceedings of the 3rd International Conference on Shellfish Restoration, Cork, Ireland, September 1999. Hydrobiologia 465 (1 – 3) 85 – 94.

26. Bell, M.V., Dick, J.R. & Kelly, M.S. 2001. Biosynthesis of eicosapentaenoic acid in the sea urchin Psammechinus miliaris (Gmelin). Lipids 36 (1) 79 – 82

27. Campbell, D.A., Kelly, M.S., Busman, M., Bolch, C.J., Wiggins, E., Moeller, P.D.R., Morton, S.L., Hess, P. and Shumway, S.E. 2001. Amnesic shellfish poisoning in the King Scallop, Pecten maximus, from the west coast of Scotland. Journal of Shellfish Research 20 (1) 75 – 84

28. Campbell, D.A., Pantazis, P. and Kelly, M.S. 2001. Impact and Residence Time of Oxytetracycline in the sea urchin Psammechinus miliaris, A Potential Aquaculture Species. Aquaculture 202 (1-2) 73 – 87

29. McLaughlin, G and Kelly, M.S. 2001. Effect of Artificial Diets Containing Carotenoid-Rich Microalgae on Gonad Growth and Colour in the sea urchin Psammechinus miliaris (Gmelin). Journal of Shellfish Research 20 (1) 377 – 382

30. Kelly, M.S. 2001. Environmental parameters controlling gametogenesis in the echinoid Psammechinus miliaris. Journal of Experimental Marine Biology and Ecology 266 (1) 67 – 80

31. Kelly, M.S. and Cook, E. J. 2001. The ecology of Psammechinus miliaris. In The Biology and Ecology of Edible Echinoid Species (J. Lawrence, Editor) Elsevier Science, Amsterdam. Pp 217 – 224

32. Kelly, M.S. 2000. The reproductive cycle of the sea urchin Psammechinus miliaris (Echinodermata: Echinoidea) in a Scottish sea loch. Journal of the Marine Biological Association 80, 909 – 919.

33. Kelly, M.S. Hunter, A.J., Scholfield, C. and McKenzie, J.D. 2000. Morphology and survivorship of larval Psammechinus miliaris in response to varying food conditions. Aquaculture 183, 233 – 240

34. McKenzie, JD, Black, KD, Kelly, M.S., Newton, LS, Handley, LL, Scrimgeour, CM, Raven, JA, Henderson, RJ. 2000. Investigations into the dietary contribution of sub-cuticular bacteria to brittlestars (Ophiuroidea: Echinodermata) from Oban Bay, Scotland. J. Mar Biol Ass UK. 80, 311 – 320

35. Cook, E.J., Bell, M.V., Black, K.D. and Kelly, M.S. 2000. Fatty acid compositions of gonadal material and diets of the sea urchin Psammechinus miliaris: Trophic and Nutritional Implications. J. Exp. Mar. Biol. Ecol. 255 (2) 261 – 274

36. Pantazis, P., Black, K.D., Connolly, J.G. and Kelly, M.S. 2000. The effect of artificial diets on growth, lipid utilisation and gonad biochemistry in the adult sea urchin Psammechinus miliaris. Journal of Shellfish Research: 19 (2) 995 – 1002

37. Kelly, M.S, Brodie, C.C and McKenzie, J.D. 1998. Somatic and gonadal growth of the sea urchin Psammechinus miliaris maintained in polyculture with the Atlantic Salmon. Journal of Shellfish Research 17 (5) 1557 – 1562

38. Cook, E.J., Kelly, M.S. and McKenzie, J.D. 1998. Gonad Development and Somatic Growth in the green sea urchin, Psammechinus miliaris fed Artificial and Natural Diets. Journal of Shellfish Research 17 (5) 1549 – 1555

39. Kelly, M.S., Brodie, C.C. and McKenzie, J. D. 1998. Sea urchins in polyculture:- the way to enhanced gonad growth? Echinoderms: San Francisco: Proceedings of the 9th International Echinoderm Conference, San Francisco, 1996. Eds Mooi, R. and Telford, M. pp 707-71

40. Burnett, W.J., McKenzie, J.D. and Kelly, M.S. 1998. Co-evolution of ophiuroids and their bacterial symbionts: studies using 16s rRNA. Echinoderms: San Francisco: Proceedings of the 9th International Echinoderm Conference, San Francisco, 1996. Eds Mooi, R. and Telford, M. pp 53-59

41. Kelly, M.S. and McKenzie, J.D. 1995. A survey of the occurrence and morphology of sub-cuticular bacteria in shelf echinoderms from the north-east Atlantic. Marine Biology 123 741-756.

42. Kelly, M.S., Barker, M.F., McKenzie, J.D. and Powell, J. 1995. The Incidence and Morphology of sub-cuticular bacteria in the Echinoderm Fauna of New Zealand. Biological Bulletin 189 91-105

43. McKenzie, J.D. and Kelly, M.S. 1994. A comparison of sub-cuticular bacteria in Brittlestars (Echinodermata: Ophiuroidea) Marine Biology 120 65-80.

44. Kelly, M.S. & McKenzie, J.D. 1991. The quantification of sub-cuticular bacteria in Echinoderms. In Echinoderm Research 1991. Eds Scalera-Liaci and Cannicatti, Balkema, Rotterdam. pp 33 – 38

45. Kelly, M.S. 1990. The biological control of leatherjackets. Proceedings of a meeting on Biological Control of Pests and Diseases. Eds A. McCracken, and P. Mercer. September 1990. Agriculture and Food Science Centre, Department of Agriculture for Northern Ireland. pp 42 - 48. ISBN 0-85389-370-5.

46. Kelly, M.S. 1989. Studies on the enemies of Tipulidae (Diptera: Nematocera) in Northern Ireland. PhD thesis. The Queen’s University of Belfast.


Published Abstracts:
1. M.S. Kelly, K.D. Black, J.C. Sanderson and E.J. 2005. Cook Can we integrate to acculmulate? Utilising ‘waste’ in open water systems. Published Abstracts, European Aquaculture Society, Trondheim, August 2005.

2. Young A.J., Philip, D., Waring, C and Kelly, M.S. 2004. Carotenoid profiles of the edible echinoid Paracentrotus lividus. In J Lawrence and O. Guzman (eds) Sea urchins: fisheries and aquaculture. Pennsylvania, DEStech publications pp 172.

3. Otero, M. Kelly, M.S. and Burnell, G. 2004. Sea urchin foraging: the selection of mussel and barnacle prey is influenced by their size and abundance. In J Lawrence and O. Guzman (eds) Sea urchins: fisheries and aquaculture. Pennsylvania, DEStech publications pp 131.

4. Otero, M. Kelly, M.S. and Burnell, G. 2004. Constructing an energy budget for the sea urchins in culture: how doeos diet influence energy partitioning? In J Lawrence and O. Guzman (eds) Sea urchins: fisheries and aquaculture. Pennsylvania, DEStech publications pp 274.

5. Cousins, S.J. Gallacher, S. Kelly, M.S., Eddy, F.B., Taylor, A. and Smith, E. 2003. Effects of storage on amnesic shellfish toxin (AST) in king scallops (Pecten maximus). Comparative Biochemistry and Physiology Part A 134 S1 – S237.

6. Kelly, M.S. 2000. Environmental parameters controlling gametogenesis in the echinoid Psammechinus miliaris. In: Echinoderms 2000. Ed Barker MF Swets & Zeitlinger, Lisse, ISBN 90 5809 189 9.

7. Barker, M.F. & Kelly, M.S. 1994. The occurrence and transmission of sub-cuticular bacteria in Echinoderm larvae. In Echinoderms through Time, Echinoderms, Dijon. Ed David, B., Balkema, Rotterdam. (Abstract)

8. Cook, E.J., Kelly, M.S. and McKenzie, J.D. 1998. The effect of stocking density on the growth (test and gonadal) of Psammechinus miliaris (Gmelin) fed salmon feed. Proceedings of 5th European Conference on Echinoderms, Milan, Italy. Balkema, Rotterdam. Eds Candia Carnavli and Bonasoro. pp 146 (Abstract)

9. Kelly, M.S, McKenzie, J.D. and Barker, M.F. 1994. Sub-cuticular bacteria: Their incidence in the Echinoderms of Great Britain and New Zealand. In Echinoderms through Time, Echinoderms, Dijon. Ed David, B,Balkema, Rotterdam. pp 13 (Abstract)


Educational and career history:

PhD Agricultural Entomology     (1989 Queen's University of Belfast)

BSc 2(1) Zoology                     (1985 Queen's Univeristy of Belfast)